基因改造人千春和改造人真希奈因违反银河道路交通法被捕。 与此同时，同样是基因改造人的茜和奏，以及改造人库尔特和麦克斯也被警方抓获。警官凉子将这几对性格迥异的组合召集起来，并指派他们执行一项社区服务任务：清洁星际列车“银河☆地铁”。原本看似 简单的任务却突然发生了意想不到的转变——“银河☆地铁”失控了！ 列车上的乘务员们惊慌失措。凉子和她的同事浅见尝试了各种方法来解决问题，但很快，她们就发现自己卷入了一场重大事故……！
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